←
Proto-Germanic *uwwalǭ (inherited)
←
Proto-Germanic *uwwǭ (mention)
←
Proto-Germanic *ūfaz (mention)
←
Proto-Germanic *ūfǭ (mention)
←
Old Swedish uggla (inherited)
←
Proto-Indo-European *up- (derived)
←
Icelandic ugla (cognate)
←
Old Norse ugla (inherited)
→
Finnish ukuli (borrowed)
→
Icelandic ugla (cognate)